In memoriam

Here we remember the scholars and teachers of Yale who have enriched us all through their pursuit of light and truth.

To submit an obituary or remembrance, please send link or text to

Graeme Berlyn, professor of forest management and physiology of trees

Berlyn, a world-renowned expert on the anatomy and physiology of plants and trees who taught at YSE for more than 60 years, died Feb. 16. He was 90.

Leander E. Keck, 1928–2024

Internationally recognized New Testament scholar and former Yale Divinity School dean Leander E. Keck ’57 Ph.D. died earlier this week at age 95.

Frank Ryan, former Yale Athletics Director, dies at 87

A highly successful NFL quarterback, Ryan also earned a Ph.D. in mathematics and worked for Congress.

Willie Ruff, legendary musician and scholar who studied and taught at Yale

Willie Ruff, a renowned jazz musician and scholar who connected the Yale and New Haven communities to "America's musical art form," died on Dec. 24. He was 92.

Brett Lindenbach, PhD, 1968–2023

Lindenbach, associate professor of microbial pathogenesis and of comparative medicine at Yale School of Medicine, died on Dec. 16. He was 55.